1001 Spikes (Wii U)

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Character Colors

You can adjust your character's color palette once you collect four golden skulls.

Commander Video

To unlock Commander Video, collect 6 golden skulls.

Pokemon Reference

After you clear the first world, a long cutscene follows. Aban will talk about the temple of Poko-mum and mistakenly refer to it as Pocket Monster, which was the original name for Pokemon.

Tower of Nanar

To unlock the cooperative mode, Tower of Nanar, collect 5 golden skulls.